The Classification of Vireya Rhododendrons
A member of the botanical family Ericaceae, the genus
Rhododendron is currently home to over 900 species,
ranging from the magnificent Rhododendron rex of
western China, with leaves up to 37cm in length, to the miniature
Rhododendron caespitosum from New Guinea whose leaves
do not exceed 0.6cm. To bring order to such a large number
of diverse species, a botanical hierarchy is used to divide
the genus into smaller botanical units and sub-units based
on similarities in morphological characteristics, i.e. species
that display similar leaf and/or flower characters are grouped
together.
At the top of the Rhododendron hierarchy is the
rank of genus and this is divided into nine subgenera, one
of which contains a group of over 300 species from south-east
Asia known collectively as Subgenus Vireya. The principal
botanical characteristics that differentiate members of Subgenus
Vireya from the other subgenera of Rhododendron
include the presence of large idioblast cells within the leaves;
distinct ovary, ovule and mega-gametophyte characters and
so-called "tailed" seeds. The geographical distribution
of vireya species, centred on south-east Asia, is another
unifying factor (see distribution
map). There are also a number of negative indicators that
are useful when separating vireyas from other rhododendron
species: there is no rachis (axis) in the inflorescence -
the flowers are carried in umbels, vireya flowers contain
no blue pigment and are never spotted.
As can be seen from the diagram below, Subgenus Vireya is
sub-divided into seven Sections, with one of these Sections
further refined into five Subsections. As before, the distinctive
botanical characteristics displayed by the species within
Subgenus Vireya are used to classify to which Section
they belong. There are many different characteristics used
by taxonomists to define each grouping - from the shape and
size of the flowers to the type of scales found on the leaves
and stems. These scales are in fact multicellular hairs, most
evident to the naked eye as a rough brownish coating on the
surface of new leaves.
Extract of Vireya classification
within Rhododendron
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Genus
Rhododendron |
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I |
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Subgenus
Vireya |
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I |
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Section
Albovireya |
Section
Discovireya |
Section
Euvireya |
Section
Malayovireya |
Section
Phaeovireya |
Section
Pseudovireya |
Section
Siphonovireya |
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Subsection
Euvireya |
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Subsection
Linnaeopsis |
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Subsection
Malesia |
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Subsection
Saxifragoidea |
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Subsection
Solenovireya |
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Dr. George Argent of the Royal Botanic Garden Edinburgh, in
his book 'Rhododendrons of Subgenus Vireya', provides
comprehensive descriptions of all the known vireya species
together with detailed information on the characters of taxonomic
value used in their classification.
Argent's account of Vireya is based on morphology
(i.e. the physical characteristics exhibited), as outlined
above. However, research is currently being undertaken at
the molecular level which should eventually lead to an alternative
classification based on the phylogeny (i.e. evolutionary relationships)
of Vireya.
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